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Updated November 2019
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Protein Info
Short Name:
QRICH2
Full Name:
Glutamine-rich protein 2
Alias:
Type:
Mass (Da):
180827
Number AA:
1663
UniProt ID:
Q9H0J4
International Prot ID:
Sequence:
Retrieve full protein sequence
GO Terms
External Links
Internal Links
Cellular Component:
Uniprot
OncoNet
Molecular Function:
PhosphoSite+
KinaseNET
Biological Process:
Phosida
TranscriptoNet
STRING
Kinexus Products
Info Box
A variety of parameters were considered in the selection of putative P-sites. Confirmed P-Sites have lower Hydrophobicity Scores. The P-site Similarity Score is lower the more that it resembles typical confirmed corresponding P-Ser, P-Thr or P-Tyr sites. The Maximum KInase Score provides the calculated score for highest match of 500 human protein kinases for the amino acid sequence surrounding the target P-site as determined with Kinase Substrate Predictor V2. The Sum KInase Score provides the additive sum of the positive individual Kinase Substrate Predictor V2 scores from 500 human protein kinases. The Conservation Score is the average of the percent similarity of the human P-site with the equivalent P-site in 20 other diverse species. Click the coloured buttons below to retrieve this and other information in the Info Box, or click the orange buttons for relevant links.
Phosphosites
-7
-6
-5
-4
-3
-2
-1
0
1
2
3
4
5
6
7
Expt. conf.
Effect
Kinase
PPase
Kinexus Products
Ref.
Evol.
Kinase Pred.
P-site Match
Site 1
S9
K
D
A
A
E
E
L
S
F
A
R
V
L
L
Q
Site 2
S40
E
L
V
S
R
K
L
S
L
V
P
G
A
E
E
Site 3
T61
E
E
L
E
Q
A
I
T
D
G
W
R
A
S
Q
Site 4
S67
I
T
D
G
W
R
A
S
Q
A
G
S
E
T
L
Site 5
S71
W
R
A
S
Q
A
G
S
E
T
L
M
G
F
S
Site 6
T73
A
S
Q
A
G
S
E
T
L
M
G
F
S
K
H
Site 7
S78
S
E
T
L
M
G
F
S
K
H
G
G
F
T
S
Site 8
S85
S
K
H
G
G
F
T
S
L
T
S
P
E
G
T
Site 9
T87
H
G
G
F
T
S
L
T
S
P
E
G
T
L
S
Site 10
S88
G
G
F
T
S
L
T
S
P
E
G
T
L
S
G
Site 11
T92
S
L
T
S
P
E
G
T
L
S
G
D
S
T
K
Site 12
S94
T
S
P
E
G
T
L
S
G
D
S
T
K
Q
P
Site 13
T98
G
T
L
S
G
D
S
T
K
Q
P
S
I
E
Q
Site 14
S109
S
I
E
Q
A
L
D
S
A
S
G
L
G
P
D
Site 15
S111
E
Q
A
L
D
S
A
S
G
L
G
P
D
R
T
Site 16
T118
S
G
L
G
P
D
R
T
A
S
G
S
G
G
T
Site 17
S120
L
G
P
D
R
T
A
S
G
S
G
G
T
A
H
Site 18
S122
P
D
R
T
A
S
G
S
G
G
T
A
H
P
S
Site 19
S129
S
G
G
T
A
H
P
S
D
G
V
S
S
R
E
Site 20
S133
A
H
P
S
D
G
V
S
S
R
E
Q
S
K
V
Site 21
S134
H
P
S
D
G
V
S
S
R
E
Q
S
K
V
P
Site 22
S138
G
V
S
S
R
E
Q
S
K
V
P
S
G
T
G
Site 23
S142
R
E
Q
S
K
V
P
S
G
T
G
R
Q
Q
Q
Site 24
S165
V
P
R
L
H
Q
S
S
T
F
Q
F
K
S
D
Site 25
T166
P
R
L
H
Q
S
S
T
F
Q
F
K
S
D
S
Site 26
S171
S
S
T
F
Q
F
K
S
D
S
D
R
H
R
S
Site 27
S173
T
F
Q
F
K
S
D
S
D
R
H
R
S
R
E
Site 28
S178
S
D
S
D
R
H
R
S
R
E
K
L
T
S
T
Site 29
S184
R
S
R
E
K
L
T
S
T
Q
P
R
R
N
A
Site 30
T185
S
R
E
K
L
T
S
T
Q
P
R
R
N
A
R
Site 31
S208
P
L
A
R
D
Q
P
S
S
V
P
A
S
Q
S
Site 32
S209
L
A
R
D
Q
P
S
S
V
P
A
S
Q
S
Q
Site 33
S213
Q
P
S
S
V
P
A
S
Q
S
Q
V
H
L
R
Site 34
S215
S
S
V
P
A
S
Q
S
Q
V
H
L
R
P
D
Site 35
T229
D
R
R
G
L
E
P
T
G
M
N
Q
P
G
L
Site 36
S280
E
Q
R
M
L
P
P
S
V
P
G
R
D
Q
Q
Site 37
T294
Q
G
L
E
L
P
S
T
D
Q
H
G
L
V
S
Site 38
S301
T
D
Q
H
G
L
V
S
V
S
A
Y
Q
H
G
Site 39
T310
S
A
Y
Q
H
G
M
T
F
P
G
T
D
Q
R
Site 40
T314
H
G
M
T
F
P
G
T
D
Q
R
S
M
E
P
Site 41
S318
F
P
G
T
D
Q
R
S
M
E
P
L
G
M
D
Site 42
S332
D
Q
R
G
C
V
I
S
G
M
G
Q
Q
G
L
Site 43
T350
G
I
D
Q
Q
G
L
T
L
P
V
V
D
Q
H
Site 44
S371
T
D
Q
H
G
L
V
S
P
G
L
M
P
I
S
Site 45
T392
V
Q
P
S
L
E
A
T
G
F
I
Q
P
G
T
Site 46
T399
T
G
F
I
Q
P
G
T
E
Q
H
D
L
I
Q
Site 47
S407
E
Q
H
D
L
I
Q
S
G
R
F
Q
R
A
L
Site 48
Y420
A
L
V
Q
R
G
A
Y
Q
P
G
L
V
Q
P
Site 49
S442
V
R
P
G
M
D
Q
S
G
L
A
Q
P
G
A
Site 50
S462
V
W
P
G
M
D
Q
S
G
L
A
Q
P
G
R
Site 51
T479
H
G
L
I
Q
P
G
T
G
Q
H
D
L
V
Q
Site 52
S487
G
Q
H
D
L
V
Q
S
G
T
G
Q
G
V
L
Site 53
S547
D
Q
H
G
L
V
Q
S
G
A
D
Q
S
D
L
Site 54
S552
V
Q
S
G
A
D
Q
S
D
L
A
Q
P
G
A
Site 55
S624
Q
P
G
E
V
Q
R
S
L
V
Q
P
G
I
V
Site 56
Y781
G
L
V
Q
P
G
A
Y
L
H
D
L
S
Q
S
Site 57
S786
G
A
Y
L
H
D
L
S
Q
S
G
T
Y
P
R
Site 58
T790
H
D
L
S
Q
S
G
T
Y
P
R
G
L
V
Q
Site 59
Y791
D
L
S
Q
S
G
T
Y
P
R
G
L
V
Q
P
Site 60
Y803
V
Q
P
G
M
D
Q
Y
G
L
R
Q
P
G
A
Site 61
Y811
G
L
R
Q
P
G
A
Y
Q
P
G
L
I
A
P
Site 62
T820
P
G
L
I
A
P
G
T
K
L
R
G
S
S
T
Site 63
S825
P
G
T
K
L
R
G
S
S
T
F
Q
A
D
S
Site 64
S826
G
T
K
L
R
G
S
S
T
F
Q
A
D
S
T
Site 65
T827
T
K
L
R
G
S
S
T
F
Q
A
D
S
T
G
Site 66
S832
S
S
T
F
Q
A
D
S
T
G
F
I
S
V
R
Site 67
T833
S
T
F
Q
A
D
S
T
G
F
I
S
V
R
P
Site 68
S837
A
D
S
T
G
F
I
S
V
R
P
Y
Q
H
G
Site 69
Y841
G
F
I
S
V
R
P
Y
Q
H
G
M
V
P
P
Site 70
Y853
V
P
P
G
R
E
Q
Y
G
Q
V
S
P
L
L
Site 71
S857
R
E
Q
Y
G
Q
V
S
P
L
L
A
S
Q
G
Site 72
S867
L
A
S
Q
G
L
A
S
P
G
I
D
R
R
S
Site 73
S874
S
P
G
I
D
R
R
S
L
V
P
P
E
T
Y
Site 74
T880
R
S
L
V
P
P
E
T
Y
Q
Q
G
L
M
H
Site 75
Y881
S
L
V
P
P
E
T
Y
Q
Q
G
L
M
H
P
Site 76
T890
Q
G
L
M
H
P
G
T
D
Q
H
S
P
I
P
Site 77
S894
H
P
G
T
D
Q
H
S
P
I
P
L
S
T
G
Site 78
S899
Q
H
S
P
I
P
L
S
T
G
L
G
S
T
H
Site 79
T900
H
S
P
I
P
L
S
T
G
L
G
S
T
H
P
Site 80
S914
P
D
Q
Q
H
V
A
S
P
G
P
G
E
H
D
Site 81
Y924
P
G
E
H
D
Q
V
Y
P
D
A
A
Q
H
G
Site 82
S935
A
Q
H
G
H
A
F
S
L
F
D
S
H
D
S
Site 83
S939
H
A
F
S
L
F
D
S
H
D
S
M
Y
P
G
Site 84
S942
S
L
F
D
S
H
D
S
M
Y
P
G
Y
R
G
Site 85
Y944
F
D
S
H
D
S
M
Y
P
G
Y
R
G
P
G
Site 86
Y947
H
D
S
M
Y
P
G
Y
R
G
P
G
Y
L
S
Site 87
Y952
P
G
Y
R
G
P
G
Y
L
S
A
D
Q
H
G
Site 88
T968
E
G
L
D
P
N
R
T
R
A
S
D
R
H
G
Site 89
S971
D
P
N
R
T
R
A
S
D
R
H
G
I
P
A
Site 90
T987
K
A
P
G
Q
D
V
T
L
F
R
S
P
D
S
Site 91
S991
Q
D
V
T
L
F
R
S
P
D
S
V
D
R
V
Site 92
S994
T
L
F
R
S
P
D
S
V
D
R
V
L
S
E
Site 93
S1000
D
S
V
D
R
V
L
S
E
G
S
E
V
S
S
Site 94
S1003
D
R
V
L
S
E
G
S
E
V
S
S
E
V
L
Site 95
S1006
L
S
E
G
S
E
V
S
S
E
V
L
S
E
R
Site 96
S1007
S
E
G
S
E
V
S
S
E
V
L
S
E
R
R
Site 97
S1011
E
V
S
S
E
V
L
S
E
R
R
N
S
L
R
Site 98
S1016
V
L
S
E
R
R
N
S
L
R
R
M
S
S
S
Site 99
S1021
R
N
S
L
R
R
M
S
S
S
F
P
T
A
V
Site 100
S1022
N
S
L
R
R
M
S
S
S
F
P
T
A
V
E
Site 101
S1023
S
L
R
R
M
S
S
S
F
P
T
A
V
E
T
Site 102
T1026
R
M
S
S
S
F
P
T
A
V
E
T
F
H
L
Site 103
T1086
E
L
Q
E
Q
L
K
T
V
K
T
L
A
K
E
Site 104
T1089
E
Q
L
K
T
V
K
T
L
A
K
E
V
W
Q
Site 105
S1149
E
L
A
E
L
R
E
S
Q
D
R
G
K
A
A
Site 106
S1160
G
K
A
A
M
E
N
S
V
S
E
A
S
L
Y
Site 107
S1162
A
A
M
E
N
S
V
S
E
A
S
L
Y
L
Q
Site 108
Y1167
S
V
S
E
A
S
L
Y
L
Q
D
Q
L
D
K
Site 109
S1191
T
S
S
S
T
L
L
S
M
S
M
A
P
H
K
Site 110
S1193
S
S
T
L
L
S
M
S
M
A
P
H
K
A
H
Site 111
S1225
L
D
V
S
H
Q
V
S
T
L
V
R
R
Y
E
Site 112
Y1231
V
S
T
L
V
R
R
Y
E
Q
L
Q
D
M
V
Site 113
S1240
Q
L
Q
D
M
V
N
S
L
A
V
S
R
P
S
Site 114
S1244
M
V
N
S
L
A
V
S
R
P
S
K
K
A
K
Site 115
S1247
S
L
A
V
S
R
P
S
K
K
A
K
L
Q
R
Site 116
S1282
E
K
L
N
I
T
T
S
N
L
I
E
D
H
R
Site 117
Y1299
Q
K
D
I
A
M
L
Y
Q
G
L
E
K
L
E
Site 118
S1326
I
D
V
K
A
D
K
S
A
L
A
T
K
V
S
Site 119
T1330
A
D
K
S
A
L
A
T
K
V
S
R
V
Q
F
Site 120
S1333
S
A
L
A
T
K
V
S
R
V
Q
F
D
A
T
Site 121
T1340
S
R
V
Q
F
D
A
T
T
E
Q
L
N
H
M
Site 122
T1371
K
M
L
D
R
L
L
T
E
M
D
N
K
L
D
Site 123
S1395
L
L
E
D
R
W
K
S
L
R
Q
Q
L
R
E
Site 124
Y1407
L
R
E
R
P
P
L
Y
Q
A
D
E
A
A
A
Site 125
S1427
L
A
H
F
H
C
L
S
C
D
R
P
L
E
T
Site 126
T1434
S
C
D
R
P
L
E
T
P
V
T
G
H
A
I
Site 127
T1437
R
P
L
E
T
P
V
T
G
H
A
I
P
V
T
Site 128
S1455
P
G
L
P
G
H
H
S
I
R
P
Y
T
V
F
Site 129
Y1459
G
H
H
S
I
R
P
Y
T
V
F
E
L
E
Q
Site 130
T1460
H
H
S
I
R
P
Y
T
V
F
E
L
E
Q
V
Site 131
S1471
L
E
Q
V
R
Q
H
S
R
N
L
K
L
G
S
Site 132
S1478
S
R
N
L
K
L
G
S
A
F
P
R
G
D
L
Site 133
S1491
D
L
A
Q
M
E
Q
S
V
G
R
L
R
S
M
Site 134
S1497
Q
S
V
G
R
L
R
S
M
H
S
K
M
L
M
Site 135
S1500
G
R
L
R
S
M
H
S
K
M
L
M
N
I
E
Site 136
S1516
V
Q
I
H
F
G
G
S
T
K
A
S
S
Q
I
Site 137
T1517
Q
I
H
F
G
G
S
T
K
A
S
S
Q
I
I
Site 138
S1520
F
G
G
S
T
K
A
S
S
Q
I
I
R
E
L
Site 139
S1521
G
G
S
T
K
A
S
S
Q
I
I
R
E
L
L
Site 140
S1535
L
H
A
Q
C
L
G
S
P
C
Y
K
R
V
T
Site 141
Y1538
Q
C
L
G
S
P
C
Y
K
R
V
T
D
M
A
Site 142
T1542
S
P
C
Y
K
R
V
T
D
M
A
D
Y
T
Y
Site 143
Y1547
R
V
T
D
M
A
D
Y
T
Y
S
T
V
P
R
Site 144
T1548
V
T
D
M
A
D
Y
T
Y
S
T
V
P
R
R
Site 145
Y1549
T
D
M
A
D
Y
T
Y
S
T
V
P
R
R
C
Site 146
T1551
M
A
D
Y
T
Y
S
T
V
P
R
R
C
G
G
Site 147
S1559
V
P
R
R
C
G
G
S
H
T
L
T
Y
P
Y
Site 148
T1561
R
R
C
G
G
S
H
T
L
T
Y
P
Y
H
R
Site 149
T1563
C
G
G
S
H
T
L
T
Y
P
Y
H
R
S
R
Site 150
Y1564
G
G
S
H
T
L
T
Y
P
Y
H
R
S
R
P
Site 151
Y1566
S
H
T
L
T
Y
P
Y
H
R
S
R
P
Q
H
Site 152
S1569
L
T
Y
P
Y
H
R
S
R
P
Q
H
L
P
R
Site 153
Y1579
Q
H
L
P
R
G
L
Y
P
T
E
E
I
Q
I
Site 154
T1581
L
P
R
G
L
Y
P
T
E
E
I
Q
I
A
M
Site 155
Y1603
L
G
L
D
G
H
I
Y
K
G
R
M
D
T
R
Site 156
T1609
I
Y
K
G
R
M
D
T
R
L
P
G
I
L
R
Site 157
S1619
P
G
I
L
R
K
D
S
S
G
T
S
K
R
K
Site 158
S1620
G
I
L
R
K
D
S
S
G
T
S
K
R
K
S
Site 159
S1623
R
K
D
S
S
G
T
S
K
R
K
S
Q
Q
P
Site 160
S1627
S
G
T
S
K
R
K
S
Q
Q
P
R
P
H
V
Site 161
S1639
P
H
V
H
R
P
P
S
L
S
S
N
G
Q
L
Site 162
S1641
V
H
R
P
P
S
L
S
S
N
G
Q
L
P
S
Site 163
S1642
H
R
P
P
S
L
S
S
N
G
Q
L
P
S
R
Site 164
S1648
S
S
N
G
Q
L
P
S
R
P
Q
S
A
Q
I
Site 165
S1652
Q
L
P
S
R
P
Q
S
A
Q
I
S
A
G
N
Site 166
S1656
R
P
Q
S
A
Q
I
S
A
G
N
T
S
E
R
Legend
Confirmed in mammals
Confirmed in related proteins or other species
Predicted by Kinexus P-Site Prediction algorithm
No data/link available
Link available
Products available
2019 Kinexus Bioinformatics Corporation