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Search by protein name, UniProt number, IPI number, or 15 AA P-site sequence.
Updated November 2019
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Protein Info
Short Name:
MUC16
Full Name:
Mucin-16
Alias:
CA125; CA-125; FLJ14303; MUC-16; Mucin 16, cell surface associated; Ovarian cancer-related tumor marker CA125; Ovarian carcinoma antigen CA125
Type:
Mass (Da):
2353428
Number AA:
22152
UniProt ID:
Q8WXI7
International Prot ID:
IPI00103552
Sequence:
Retrieve full protein sequence
GO Terms
External Links
Internal Links
Cellular Component:
GO:0005615
GO:0019898
GO:0016021
Uniprot
OncoNet
Molecular Function:
GO:0005515
PhosphoSite+
KinaseNET
Biological Process:
GO:0007155
Phosida
TranscriptoNet
STRING
Kinexus Products
Info Box
A variety of parameters were considered in the selection of putative P-sites. Confirmed P-Sites have lower Hydrophobicity Scores. The P-site Similarity Score is lower the more that it resembles typical confirmed corresponding P-Ser, P-Thr or P-Tyr sites. The Maximum KInase Score provides the calculated score for highest match of 500 human protein kinases for the amino acid sequence surrounding the target P-site as determined with Kinase Substrate Predictor V2. The Sum KInase Score provides the additive sum of the positive individual Kinase Substrate Predictor V2 scores from 500 human protein kinases. The Conservation Score is the average of the percent similarity of the human P-site with the equivalent P-site in 20 other diverse species. Click the coloured buttons below to retrieve this and other information in the Info Box, or click the orange buttons for relevant links.
Phosphosites
-7
-6
-5
-4
-3
-2
-1
0
1
2
3
4
5
6
7
Expt. conf.
Effect
Kinase
PPase
Kinexus Products
Ref.
Evol.
Kinase Pred.
P-site Match
Site 1
S5
_
_
_
M
L
K
P
S
G
L
P
G
S
S
S
Site 2
T27
G
S
R
S
T
K
A
T
P
E
M
D
S
G
L
Site 3
T54
A
I
V
V
T
E
H
T
L
P
F
T
S
P
D
Site 4
T63
P
F
T
S
P
D
K
T
L
A
S
P
T
S
S
Site 5
S66
S
P
D
K
T
L
A
S
P
T
S
S
V
V
G
Site 6
S69
K
T
L
A
S
P
T
S
S
V
V
G
R
T
T
Site 7
S84
Q
S
L
G
V
M
S
S
A
L
P
E
S
T
S
Site 8
T95
E
S
T
S
R
G
M
T
H
S
E
Q
R
T
S
Site 9
S97
T
S
R
G
M
T
H
S
E
Q
R
T
S
P
S
Site 10
S127
T
S
M
V
S
G
L
S
S
P
R
T
R
T
S
Site 11
S128
S
M
V
S
G
L
S
S
P
R
T
R
T
S
S
Site 12
Y162
S
G
P
V
T
E
K
Y
T
V
P
T
E
T
S
Site 13
T163
G
P
V
T
E
K
Y
T
V
P
T
E
T
S
T
Site 14
T166
T
E
K
Y
T
V
P
T
E
T
S
T
T
E
G
Site 15
S202
T
F
A
D
S
T
A
S
K
E
N
A
P
V
S
Site 16
T211
E
N
A
P
V
S
M
T
P
A
E
T
T
V
T
Site 17
T222
T
T
V
T
D
S
H
T
P
G
R
T
N
P
S
Site 18
S229
T
P
G
R
T
N
P
S
F
G
T
L
Y
S
S
Site 19
S299
D
N
K
I
S
E
T
S
I
F
S
G
Q
S
L
Site 20
S311
Q
S
L
T
S
P
L
S
P
G
V
P
E
A
R
Site 21
S354
I
S
S
L
G
T
P
S
I
S
T
K
Q
T
A
Site 22
S475
I
R
S
P
S
Q
V
S
S
S
H
P
T
R
E
Site 23
T486
P
T
R
E
L
R
T
T
G
S
T
S
G
R
Q
Site 24
S494
G
S
T
S
G
R
Q
S
S
S
T
A
A
H
G
Site 25
S495
S
T
S
G
R
Q
S
S
S
T
A
A
H
G
S
Site 26
S496
T
S
G
R
Q
S
S
S
T
A
A
H
G
S
S
Site 27
T497
S
G
R
Q
S
S
S
T
A
A
H
G
S
S
D
Site 28
S502
S
S
T
A
A
H
G
S
S
D
I
L
R
A
T
Site 29
S503
S
T
A
A
H
G
S
S
D
I
L
R
A
T
T
Site 30
S511
D
I
L
R
A
T
T
S
S
T
S
K
A
S
S
Site 31
S708
L
N
T
R
H
P
F
S
S
P
E
P
D
S
A
Site 32
S709
N
T
R
H
P
F
S
S
P
E
P
D
S
A
G
Site 33
T718
E
P
D
S
A
G
H
T
K
I
S
T
S
I
P
Site 34
S844
T
L
S
L
P
S
V
S
G
V
K
T
T
F
S
Site 35
S890
R
V
R
T
T
L
A
S
T
S
V
P
T
P
V
Site 36
T895
L
A
S
T
S
V
P
T
P
V
F
P
T
M
D
Site 37
S927
L
R
A
T
S
S
T
S
V
T
N
S
T
G
S
Site 38
T932
S
T
S
V
T
N
S
T
G
S
A
L
P
K
I
Site 39
T955
M
S
Q
T
N
R
D
T
F
N
D
S
A
A
P
Site 40
S959
N
R
D
T
F
N
D
S
A
A
P
Q
S
T
T
Site 41
T984
G
L
P
S
A
T
T
T
V
S
T
S
A
T
S
Site 42
T1002
T
V
M
V
S
K
F
T
S
P
A
T
S
S
M
Site 43
S1007
K
F
T
S
P
A
T
S
S
M
E
A
T
S
I
Site 44
T1202
L
P
S
T
T
S
L
T
P
S
N
I
P
T
S
Site 45
T1770
I
G
G
S
T
F
W
T
K
E
V
T
T
V
M
Site 46
S1785
T
S
D
I
S
K
S
S
A
R
T
E
S
S
S
Site 47
T1794
R
T
E
S
S
S
A
T
L
M
S
T
A
L
G
Site 48
S2007
Q
S
T
P
Y
V
N
S
V
A
V
S
A
S
P
Site 49
T2049
I
S
R
R
N
A
I
T
S
W
L
W
D
L
T
Site 50
T2067
P
T
T
T
W
P
S
T
S
L
S
E
A
L
S
Site 51
S2070
T
W
P
S
T
S
L
S
E
A
L
S
S
G
H
Site 52
T2270
I
S
H
F
P
E
K
T
T
H
S
F
R
T
I
Site 53
T2376
S
S
S
D
L
I
N
T
S
I
A
S
S
T
M
Site 54
S2381
I
N
T
S
I
A
S
S
T
M
D
V
T
K
T
Site 55
T2454
P
S
T
L
P
P
F
T
I
T
H
P
V
E
T
Site 56
T2456
T
L
P
P
F
T
I
T
H
P
V
E
T
S
S
Site 57
T2461
T
I
T
H
P
V
E
T
S
S
A
L
L
A
W
Site 58
S2798
G
S
Q
G
M
A
S
S
G
G
I
R
S
G
S
Site 59
T2820
T
F
S
S
L
P
L
T
M
N
P
G
E
V
T
Site 60
S2924
T
I
P
D
S
D
A
S
T
A
S
S
S
L
S
Site 61
T2925
I
P
D
S
D
A
S
T
A
S
S
S
L
S
K
Site 62
S2928
S
D
A
S
T
A
S
S
S
L
S
K
S
P
E
Site 63
S2992
D
P
R
Y
A
S
E
S
M
S
Y
P
D
P
S
Site 64
S2994
R
Y
A
S
E
S
M
S
Y
P
D
P
S
K
A
Site 65
S2999
S
M
S
Y
P
D
P
S
K
A
S
S
A
M
T
Site 66
S3049
S
F
L
S
P
T
A
S
T
S
R
K
T
S
L
Site 67
T3054
T
A
S
T
S
R
K
T
S
L
F
L
G
P
S
Site 68
S3061
T
S
L
F
L
G
P
S
M
A
R
Q
P
N
I
Site 69
T3096
Q
E
E
P
P
E
L
T
S
S
Q
T
I
A
E
Site 70
S3097
E
E
P
P
E
L
T
S
S
Q
T
I
A
E
E
Site 71
S3098
E
P
P
E
L
T
S
S
Q
T
I
A
E
E
E
Site 72
T3100
P
E
L
T
S
S
Q
T
I
A
E
E
E
G
T
Site 73
T3132
L
L
P
V
S
S
P
T
E
P
T
A
R
R
K
Site 74
T3359
T
G
G
T
T
G
D
T
H
V
S
L
S
T
S
Site 75
S3364
G
D
T
H
V
S
L
S
T
S
S
N
I
L
E
Site 76
S3485
P
S
G
G
K
T
S
S
A
S
S
V
T
S
P
Site 77
T3663
E
T
S
T
S
E
R
T
L
S
P
S
D
T
T
Site 78
S3665
S
T
S
E
R
T
L
S
P
S
D
T
T
A
S
Site 79
S3679
S
T
P
I
S
T
F
S
R
V
Q
R
M
S
I
Site 80
S3695
V
P
D
I
L
S
T
S
W
T
P
S
S
T
E
Site 81
S3987
T
S
P
V
V
T
T
S
T
R
E
Q
A
I
V
Site 82
S4688
T
T
T
Q
T
G
S
S
G
A
T
S
K
I
P
Site 83
T4765
L
P
S
S
V
A
F
T
P
Q
W
H
S
T
S
Site 84
T4821
S
V
T
S
A
A
T
T
D
I
E
T
T
H
P
Site 85
S4893
R
T
E
T
E
T
T
S
S
L
T
P
K
L
R
Site 86
S4971
T
S
P
I
M
T
E
S
A
E
I
T
I
T
T
Site 87
S4996
D
T
F
T
M
D
P
S
N
T
T
P
Q
A
G
Site 88
S5022
L
D
V
T
T
L
M
S
R
I
P
Q
D
V
S
Site 89
T5071
D
K
L
S
S
P
M
T
S
L
L
T
S
G
L
Site 90
S5094
T
R
L
E
P
V
T
S
S
L
P
N
F
S
S
Site 91
T5187
T
N
I
K
R
E
P
T
Y
F
L
T
P
R
L
Site 92
T5191
R
E
P
T
Y
F
L
T
P
R
L
R
E
T
S
Site 93
S5210
S
S
F
P
T
D
T
S
F
L
L
S
K
V
P
Site 94
S5467
T
L
G
D
T
S
V
S
T
S
T
P
N
I
S
Site 95
T5481
S
Q
T
N
Q
I
Q
T
E
P
T
A
S
L
S
Site 96
S5488
T
E
P
T
A
S
L
S
P
R
L
R
E
S
S
Site 97
T5524
A
I
T
Q
A
S
R
T
E
I
S
S
S
R
T
Site 98
S5527
Q
A
S
R
T
E
I
S
S
S
R
T
S
I
S
Site 99
T5531
T
E
I
S
S
S
R
T
S
I
S
D
L
D
R
Site 100
S5532
E
I
S
S
S
R
T
S
I
S
D
L
D
R
P
Site 101
S5737
S
G
H
E
S
Q
S
S
V
P
A
D
S
H
T
Site 102
S5749
S
H
T
S
K
A
T
S
P
M
G
I
T
F
A
Site 103
T5754
A
T
S
P
M
G
I
T
F
A
M
G
D
T
S
Site 104
T5845
S
P
Y
I
S
T
E
T
I
T
R
L
S
T
F
Site 105
S5850
T
E
T
I
T
R
L
S
T
F
P
F
V
T
G
Site 106
T5859
F
P
F
V
T
G
S
T
E
M
A
I
T
N
Q
Site 107
T6116
E
V
S
R
A
E
V
T
Y
Y
D
P
T
F
M
Site 108
S6332
S
D
P
Y
K
A
T
S
A
V
V
I
T
S
T
Site 109
T6337
A
T
S
A
V
V
I
T
S
T
S
G
D
S
I
Site 110
T6411
E
L
T
S
S
S
R
T
S
I
Q
G
T
E
K
Site 111
T6585
T
S
E
D
I
K
D
T
E
K
M
Y
P
S
T
Site 112
T6592
T
E
K
M
Y
P
S
T
N
I
A
V
T
N
V
Site 113
S6950
I
K
R
E
S
T
Y
S
L
T
P
G
L
R
E
Site 114
S6959
T
P
G
L
R
E
T
S
I
S
Q
N
A
S
F
Site 115
S6965
T
S
I
S
Q
N
A
S
F
S
T
D
T
S
I
Site 116
T7014
L
P
E
I
S
T
R
T
M
T
R
L
F
A
S
Site 117
T7016
E
I
S
T
R
T
M
T
R
L
F
A
S
P
T
Site 118
S7245
I
Q
K
E
S
I
S
S
L
S
P
K
L
R
E
Site 119
S7281
I
G
A
T
T
E
I
S
R
T
E
V
T
S
S
Site 120
S7306
S
T
M
L
P
E
I
S
T
T
R
K
I
I
K
Site 121
T7327
L
A
E
S
S
E
M
T
I
K
T
Q
T
S
P
Site 122
S7339
T
S
P
P
G
S
T
S
E
S
T
F
T
L
D
Site 123
S7508
S
E
T
T
K
A
T
S
A
M
G
T
I
S
I
Site 124
T7519
T
I
S
I
M
G
D
T
S
V
S
T
L
T
P
Site 125
S7520
I
S
I
M
G
D
T
S
V
S
T
L
T
P
A
Site 126
T7549
L
T
T
R
L
R
E
T
S
T
S
E
E
T
S
Site 127
S7550
T
T
R
L
R
E
T
S
T
S
E
E
T
S
L
Site 128
T7551
T
R
L
R
E
T
S
T
S
E
E
T
S
L
A
Site 129
T7706
T
S
P
H
P
V
S
T
T
F
L
A
H
I
P
Site 130
S7715
F
L
A
H
I
P
P
S
P
L
P
V
T
S
L
Site 131
S7725
P
V
T
S
L
L
T
S
G
P
A
T
T
T
D
Site 132
T7757
T
T
S
H
E
R
L
T
T
Y
K
D
T
A
H
Site 133
T7758
T
S
H
E
R
L
T
T
Y
K
D
T
A
H
T
Site 134
T7762
R
L
T
T
Y
K
D
T
A
H
T
E
A
V
H
Site 135
T7844
E
S
S
G
S
E
G
T
S
S
G
T
K
M
S
Site 136
S7855
T
K
M
S
T
V
L
S
K
V
P
T
G
A
T
Site 137
T7859
T
V
L
S
K
V
P
T
G
A
T
T
E
I
S
Site 138
S8068
T
D
V
G
T
S
S
S
G
H
E
S
T
S
F
Site 139
S8080
T
S
F
V
L
A
D
S
Q
T
S
K
V
T
S
Site 140
S8099
T
S
T
M
E
D
T
S
V
S
T
S
T
P
G
Site 141
S8111
T
P
G
F
F
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Site 142
T8122
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Site 143
T8161
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Site 144
S8162
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T8166
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Site 146
T8381
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Site 147
S8382
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S8424
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S8425
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S8426
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S8444
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T8470
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T8477
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T8481
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Site 155
T8721
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Site 156
S8723
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T8725
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Site 158
T8781
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Site 159
T8788
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H
Site 160
T8790
G
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A
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Site 161
T8805
A
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Site 162
S8825
H
S
A
A
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Site 163
S8828
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Site 164
T8831
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Site 165
S8835
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G
M
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Site 166
S8969
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S
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Site 167
S8989
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A
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M
Site 168
T8992
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P
A
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Site 169
T9295
L
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Site 170
S9296
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P
G
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Site 171
T9365
P
L
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M
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K
Site 172
T9370
E
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K
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G
P
S
G
Site 173
T9373
D
M
A
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G
P
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G
A
T
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Site 174
T9385
A
T
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Q
G
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F
T
L
D
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T
A
Site 175
T9564
I
R
D
N
M
V
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T
T
M
P
G
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S
G
Site 176
T9565
R
D
N
M
V
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T
T
M
P
G
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S
G
I
Site 177
S9569
V
S
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T
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G
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I
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Site 178
S9581
R
I
E
I
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P
G
L
R
Site 179
S9651
D
E
V
V
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L
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I
M
T
E
Site 180
T9738
S
L
T
S
L
P
L
T
T
S
L
S
P
V
S
Site 181
S9740
T
S
L
P
L
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T
S
L
S
P
V
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S
T
Site 182
S9742
L
P
L
T
T
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L
S
P
V
S
S
T
L
L
Site 183
S9761
S
S
P
L
P
V
T
S
L
I
L
P
G
L
V
Site 184
S9822
A
V
A
K
V
R
T
S
S
S
V
H
E
S
H
Site 185
S9836
H
S
S
V
L
A
D
S
E
T
T
I
T
I
P
Site 186
T9854
I
T
S
A
V
D
D
T
T
V
F
T
S
N
P
Site 187
S10145
P
S
S
M
A
E
T
S
I
S
T
S
M
P
A
Site 188
S10147
S
M
A
E
T
S
I
S
T
S
M
P
A
N
F
Site 189
T10148
M
A
E
T
S
I
S
T
S
M
P
A
N
F
E
Site 190
T10340
P
F
S
S
S
T
F
T
T
T
D
S
S
T
I
Site 191
S10344
S
T
F
T
T
T
D
S
S
T
I
P
A
L
H
Site 192
S10382
E
G
R
L
V
M
V
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T
L
D
T
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S
Q
Site 193
S10387
M
V
S
T
L
D
T
S
S
Q
P
G
R
T
S
Site 194
T10447
N
E
A
A
H
R
G
T
I
R
P
V
K
G
P
Site 195
S10462
Q
T
S
T
S
P
A
S
P
K
G
L
H
T
G
Site 196
T10468
A
S
P
K
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L
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T
G
G
T
K
R
M
E
Site 197
T10471
K
G
L
H
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G
G
T
K
R
M
E
T
T
T
Site 198
T10476
G
G
T
K
R
M
E
T
T
T
T
A
L
K
T
Site 199
T10483
T
T
T
T
A
L
K
T
T
T
T
A
L
K
T
Site 200
S10572
R
E
S
E
T
T
A
S
L
V
S
R
S
G
A
Site 201
T10778
V
T
H
P
E
A
Q
T
S
S
A
I
P
T
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Site 202
T10796
P
A
V
S
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L
V
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S
M
V
T
S
L
A
Site 203
T10806
V
T
S
L
A
A
K
T
S
T
T
N
R
A
L
Site 204
S10849
S
I
F
F
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K
S
D
T
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M
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Site 205
S10864
T
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H
G
A
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A
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P
T
Site 206
S10865
S
H
G
A
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A
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T
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Site 207
T10869
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A
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E
V
Site 208
S11093
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T
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P
G
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G
V
V
Site 209
S11097
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G
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L
V
Site 210
S11459
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A
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M
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L
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Site 211
T11463
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G
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Site 212
S11464
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A
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Site 213
S11536
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E
A
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A
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P
D
Site 214
S11550
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M
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D
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L
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G
T
Site 215
T11569
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F
P
T
L
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E
T
P
Y
E
P
E
T
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Site 216
Y11571
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L
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Y
E
P
E
T
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Site 217
T11644
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T
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A
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D
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A
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T
L
Site 218
T11650
D
T
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T
A
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L
T
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P
G
E
Site 219
T11672
A
T
H
P
G
T
Q
T
G
F
T
V
P
I
R
Site 220
S11711
S
R
T
T
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S
F
S
H
S
S
P
D
A
T
Site 221
S11747
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A
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G
A
Site 222
T11758
S
S
G
A
A
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T
T
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P
T
L
T
H
Site 223
T11762
A
T
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T
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T
L
T
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G
M
Site 224
T11772
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E
T
T
A
L
L
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T
H
Site 225
T11876
L
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L
G
L
L
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L
L
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Site 226
T11890
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A
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T
L
T
V
Site 227
T11911
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A
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Q
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V
T
Site 228
T11998
K
T
T
T
T
F
N
T
L
A
G
S
L
F
T
Site 229
S12017
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G
M
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A
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S
V
T
S
R
T
Site 230
S12019
M
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T
L
A
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V
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S
R
T
S
Y
Site 231
S12025
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S
V
T
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R
T
S
Y
N
H
R
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W
I
Site 232
S12262
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K
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N
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G
L
L
K
Site 233
S12483
E
R
L
Y
W
E
L
S
Q
L
T
N
G
I
K
Site 234
T12486
Y
W
E
L
S
Q
L
T
N
G
I
K
E
L
G
Site 235
S12639
E
Q
L
Y
W
E
L
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Q
L
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N
G
I
K
Site 236
S12781
T
H
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L
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K
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P
G
V
D
R
E
Q
Site 237
S13134
V
N
G
F
T
H
R
S
S
V
A
P
T
S
T
Site 238
T13144
A
P
T
S
T
P
G
T
S
T
V
D
L
G
T
Site 239
S13160
G
T
P
S
S
L
P
S
P
T
T
A
V
P
L
Site 240
Y13182
F
T
I
T
N
L
Q
Y
G
E
D
M
R
H
P
Site 241
Y13258
G
L
D
R
E
Q
L
Y
W
Q
L
S
Q
M
T
Site 242
S13503
E
N
M
Q
H
P
G
S
R
K
F
N
T
T
E
Site 243
Y15148
S
V
T
E
L
G
P
Y
T
L
D
R
D
S
L
Site 244
T15299
S
Q
L
T
H
G
I
T
E
L
G
P
Y
T
L
Site 245
T15421
P
K
K
D
G
A
A
T
K
V
D
A
I
C
T
Site 246
Y15429
K
V
D
A
I
C
T
Y
R
P
D
P
K
S
P
Site 247
S16213
T
H
R
V
D
P
K
S
P
G
V
D
R
E
Q
Site 248
Y20901
G
L
D
R
Q
Q
L
Y
W
E
L
S
Q
L
T
Site 249
T21881
D
I
Q
D
K
V
T
T
L
Y
K
G
S
Q
L
Site 250
T22122
L
I
C
G
V
L
V
T
T
R
R
R
K
K
E
Site 251
T22123
I
C
G
V
L
V
T
T
R
R
R
K
K
E
G
Site 252
Y22132
R
R
K
K
E
G
E
Y
N
V
Q
Q
Q
C
P
Site 253
Y22141
V
Q
Q
Q
C
P
G
Y
Y
Q
S
H
L
D
L
Site 254
Y22142
Q
Q
Q
C
P
G
Y
Y
Q
S
H
L
D
L
E
Site 255
S22144
Q
C
P
G
Y
Y
Q
S
H
L
D
L
E
D
L
Legend
Confirmed in mammals
Confirmed in related proteins or other species
Predicted by Kinexus P-Site Prediction algorithm
No data/link available
Link available
Products available
2019 Kinexus Bioinformatics Corporation