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Updated November 2019
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Protein Info
Short Name:
GOLGA4
Full Name:
Golgin subfamily A member 4
Alias:
256 kDa golgin; 72.1 protein; GCP2; GOA4; GOGA4; GOLG; Golgi autoantigen, golgin subfamily A member 4; Golgi autoantigen, golgin subfamily a, 4; Golgin 245; Golgin-240; P230; Trans-Golgi p230
Type:
Vesicle protein
Mass (Da):
261140
Number AA:
2230
UniProt ID:
Q13439
International Prot ID:
IPI00013272
Sequence:
Retrieve full protein sequence
GO Terms
External Links
Internal Links
Cellular Component:
GO:0000139
GO:0005737
GO:0005794
Uniprot
OncoNet
Molecular Function:
GO:0000166
GO:0005488
PhosphoSite+
KinaseNET
Biological Process:
GO:0006810
GO:0009987
GO:0016192
Phosida
TranscriptoNet
STRING
Kinexus Products
Info Box
A variety of parameters were considered in the selection of putative P-sites. Confirmed P-Sites have lower Hydrophobicity Scores. The P-site Similarity Score is lower the more that it resembles typical confirmed corresponding P-Ser, P-Thr or P-Tyr sites. The Maximum KInase Score provides the calculated score for highest match of 500 human protein kinases for the amino acid sequence surrounding the target P-site as determined with Kinase Substrate Predictor V2. The Sum KInase Score provides the additive sum of the positive individual Kinase Substrate Predictor V2 scores from 500 human protein kinases. The Conservation Score is the average of the percent similarity of the human P-site with the equivalent P-site in 20 other diverse species. Click the coloured buttons below to retrieve this and other information in the Info Box, or click the orange buttons for relevant links.
Phosphosites
-7
-6
-5
-4
-3
-2
-1
0
1
2
3
4
5
6
7
Expt. conf.
Effect
Kinase
PPase
Kinexus Products
Ref.
Evol.
Kinase Pred.
P-site Match
Site 1
S10
K
K
L
K
Q
K
I
S
E
E
Q
Q
Q
L
Q
Site 2
S26
A
L
A
P
A
Q
A
S
S
N
S
S
T
P
T
Site 3
S27
L
A
P
A
Q
A
S
S
N
S
S
T
P
T
R
Site 4
S29
P
A
Q
A
S
S
N
S
S
T
P
T
R
M
R
Site 5
S30
A
Q
A
S
S
N
S
S
T
P
T
R
M
R
S
Site 6
T31
Q
A
S
S
N
S
S
T
P
T
R
M
R
S
R
Site 7
T33
S
S
N
S
S
T
P
T
R
M
R
S
R
T
S
Site 8
S37
S
T
P
T
R
M
R
S
R
T
S
S
F
T
E
Site 9
T39
P
T
R
M
R
S
R
T
S
S
F
T
E
Q
L
Site 10
S40
T
R
M
R
S
R
T
S
S
F
T
E
Q
L
D
Site 11
S41
R
M
R
S
R
T
S
S
F
T
E
Q
L
D
E
Site 12
T43
R
S
R
T
S
S
F
T
E
Q
L
D
E
G
T
Site 13
T50
T
E
Q
L
D
E
G
T
P
N
R
E
S
G
D
Site 14
S55
E
G
T
P
N
R
E
S
G
D
T
Q
S
F
A
Site 15
T58
P
N
R
E
S
G
D
T
Q
S
F
A
Q
K
L
Site 16
S60
R
E
S
G
D
T
Q
S
F
A
Q
K
L
Q
L
Site 17
S71
K
L
Q
L
R
V
P
S
V
E
S
L
F
R
S
Site 18
S74
L
R
V
P
S
V
E
S
L
F
R
S
P
I
K
Site 19
S78
S
V
E
S
L
F
R
S
P
I
K
E
S
L
F
Site 20
S83
F
R
S
P
I
K
E
S
L
F
R
S
S
S
K
Site 21
S87
I
K
E
S
L
F
R
S
S
S
K
E
S
L
V
Site 22
S88
K
E
S
L
F
R
S
S
S
K
E
S
L
V
R
Site 23
S89
E
S
L
F
R
S
S
S
K
E
S
L
V
R
T
Site 24
S92
F
R
S
S
S
K
E
S
L
V
R
T
S
S
R
Site 25
T96
S
K
E
S
L
V
R
T
S
S
R
E
S
L
N
Site 26
S97
K
E
S
L
V
R
T
S
S
R
E
S
L
N
R
Site 27
S98
E
S
L
V
R
T
S
S
R
E
S
L
N
R
L
Site 28
S101
V
R
T
S
S
R
E
S
L
N
R
L
D
L
D
Site 29
S109
L
N
R
L
D
L
D
S
S
T
A
S
F
D
P
Site 30
S110
N
R
L
D
L
D
S
S
T
A
S
F
D
P
P
Site 31
T111
R
L
D
L
D
S
S
T
A
S
F
D
P
P
S
Site 32
S113
D
L
D
S
S
T
A
S
F
D
P
P
S
D
M
Site 33
S118
T
A
S
F
D
P
P
S
D
M
D
S
E
A
E
Site 34
S122
D
P
P
S
D
M
D
S
E
A
E
D
L
V
G
Site 35
S133
D
L
V
G
N
S
D
S
L
N
K
E
Q
L
I
Site 36
S149
R
L
R
R
M
E
R
S
L
S
S
Y
R
G
K
Site 37
S151
R
R
M
E
R
S
L
S
S
Y
R
G
K
Y
S
Site 38
S152
R
M
E
R
S
L
S
S
Y
R
G
K
Y
S
E
Site 39
Y153
M
E
R
S
L
S
S
Y
R
G
K
Y
S
E
L
Site 40
Y157
L
S
S
Y
R
G
K
Y
S
E
L
V
T
A
Y
Site 41
T162
G
K
Y
S
E
L
V
T
A
Y
Q
M
L
Q
R
Site 42
S179
K
K
L
Q
G
I
L
S
Q
S
Q
D
K
S
L
Site 43
S181
L
Q
G
I
L
S
Q
S
Q
D
K
S
L
R
R
Site 44
S185
L
S
Q
S
Q
D
K
S
L
R
R
I
A
E
L
Site 45
S213
L
Q
E
E
F
D
A
S
L
E
E
K
D
Q
Y
Site 46
Y220
S
L
E
E
K
D
Q
Y
I
S
V
L
Q
T
Q
Site 47
T226
Q
Y
I
S
V
L
Q
T
Q
V
S
L
L
K
Q
Site 48
S229
S
V
L
Q
T
Q
V
S
L
L
K
Q
R
L
R
Site 49
S266
T
K
E
E
N
P
E
S
D
G
E
P
V
V
E
Site 50
T276
E
P
V
V
E
D
G
T
S
V
K
T
L
E
T
Site 51
S277
P
V
V
E
D
G
T
S
V
K
T
L
E
T
L
Site 52
T280
E
D
G
T
S
V
K
T
L
E
T
L
Q
Q
R
Site 53
T342
D
L
H
M
A
E
K
T
K
L
I
T
Q
L
R
Site 54
T346
A
E
K
T
K
L
I
T
Q
L
R
D
A
K
N
Site 55
T369
K
G
M
V
I
A
E
T
K
R
Q
M
H
E
T
Site 56
T376
T
K
R
Q
M
H
E
T
L
E
M
K
E
E
E
Site 57
T395
R
S
R
I
K
Q
M
T
T
Q
G
E
E
L
R
Site 58
S408
L
R
E
Q
K
E
K
S
E
R
A
A
F
E
E
Site 59
S421
E
E
L
E
K
A
L
S
T
A
Q
K
T
E
E
Site 60
T443
E
M
D
E
Q
I
K
T
I
E
K
T
S
E
E
Site 61
S454
T
S
E
E
E
R
I
S
L
Q
Q
E
L
S
R
Site 62
S460
I
S
L
Q
Q
E
L
S
R
V
K
Q
E
V
V
Site 63
S473
V
V
D
V
M
K
K
S
S
E
E
Q
I
A
K
Site 64
T497
A
R
K
E
Q
E
L
T
K
K
L
Q
T
R
E
Site 65
S518
M
K
V
A
L
E
K
S
Q
S
E
Y
L
K
I
Site 66
S520
V
A
L
E
K
S
Q
S
E
Y
L
K
I
S
Q
Site 67
Y522
L
E
K
S
Q
S
E
Y
L
K
I
S
Q
E
K
Site 68
S526
Q
S
E
Y
L
K
I
S
Q
E
K
E
Q
Q
E
Site 69
S534
Q
E
K
E
Q
Q
E
S
L
A
L
E
E
L
E
Site 70
S551
K
K
A
I
L
T
E
S
E
N
K
L
R
D
L
Site 71
Y565
L
Q
Q
E
A
E
T
Y
R
T
R
I
L
E
L
Site 72
S574
T
R
I
L
E
L
E
S
S
L
E
K
S
L
Q
Site 73
S575
R
I
L
E
L
E
S
S
L
E
K
S
L
Q
E
Site 74
S579
L
E
S
S
L
E
K
S
L
Q
E
N
K
N
Q
Site 75
T606
N
K
H
N
K
E
I
T
V
M
V
E
K
H
K
Site 76
T614
V
M
V
E
K
H
K
T
E
L
E
S
L
K
H
Site 77
Y638
L
Q
V
L
K
Q
Q
Y
Q
T
E
M
E
K
L
Site 78
T684
E
K
L
D
V
K
Q
T
E
L
E
S
L
S
S
Site 79
S688
V
K
Q
T
E
L
E
S
L
S
S
E
L
S
E
Site 80
S690
Q
T
E
L
E
S
L
S
S
E
L
S
E
V
L
Site 81
S691
T
E
L
E
S
L
S
S
E
L
S
E
V
L
K
Site 82
S694
E
S
L
S
S
E
L
S
E
V
L
K
A
R
H
Site 83
S708
H
K
L
E
E
E
L
S
V
L
K
D
Q
T
D
Site 84
T714
L
S
V
L
K
D
Q
T
D
K
M
K
Q
E
L
Site 85
S746
I
I
K
E
H
E
V
S
I
Q
R
T
E
K
A
Site 86
S805
A
K
L
D
V
F
Q
S
Y
Q
S
A
T
H
E
Site 87
Y806
K
L
D
V
F
Q
S
Y
Q
S
A
T
H
E
Q
Site 88
T833
Q
K
L
L
D
L
E
T
E
R
I
L
L
T
K
Site 89
S875
Q
Q
L
E
K
Q
N
S
E
M
E
Q
K
V
K
Site 90
S883
E
M
E
Q
K
V
K
S
L
T
Q
V
Y
E
S
Site 91
T885
E
Q
K
V
K
S
L
T
Q
V
Y
E
S
K
L
Site 92
T902
G
N
K
E
Q
E
Q
T
K
Q
I
L
V
E
K
Site 93
T928
K
K
E
I
E
I
L
T
Q
K
L
S
A
K
E
Site 94
S932
E
I
L
T
Q
K
L
S
A
K
E
D
S
I
H
Site 95
Y945
I
H
I
L
N
E
E
Y
E
T
K
F
K
N
Q
Site 96
T969
K
A
K
E
M
Q
E
T
L
K
K
K
L
L
D
Site 97
T988
L
K
K
E
L
E
N
T
A
L
E
L
S
Q
K
Site 98
S993
E
N
T
A
L
E
L
S
Q
K
E
K
Q
F
N
Site 99
S1011
L
E
M
A
Q
A
N
S
A
G
I
S
D
A
V
Site 100
S1015
Q
A
N
S
A
G
I
S
D
A
V
S
R
L
E
Site 101
S1019
A
G
I
S
D
A
V
S
R
L
E
T
N
Q
K
Site 102
T1023
D
A
V
S
R
L
E
T
N
Q
K
E
Q
I
E
Site 103
S1031
N
Q
K
E
Q
I
E
S
L
T
E
V
H
R
R
Site 104
S1045
R
E
L
N
D
V
I
S
I
W
E
K
K
L
N
Site 105
T1107
E
G
V
K
Q
D
T
T
L
N
E
L
Q
E
Q
Site 106
S1119
Q
E
Q
L
K
Q
K
S
A
H
V
N
S
L
A
Site 107
S1146
L
E
V
D
L
N
K
S
L
K
E
N
T
F
L
Site 108
T1151
N
K
S
L
K
E
N
T
F
L
Q
E
Q
L
V
Site 109
S1172
E
E
D
K
R
K
V
S
E
L
T
S
K
L
K
Site 110
S1176
R
K
V
S
E
L
T
S
K
L
K
T
T
D
E
Site 111
T1180
E
L
T
S
K
L
K
T
T
D
E
E
F
Q
S
Site 112
S1187
T
T
D
E
E
F
Q
S
L
K
S
S
H
E
K
Site 113
S1190
E
E
F
Q
S
L
K
S
S
H
E
K
S
N
K
Site 114
S1191
E
F
Q
S
L
K
S
S
H
E
K
S
N
K
S
Site 115
S1195
L
K
S
S
H
E
K
S
N
K
S
L
E
D
K
Site 116
S1198
S
H
E
K
S
N
K
S
L
E
D
K
S
L
E
Site 117
S1203
N
K
S
L
E
D
K
S
L
E
F
K
K
L
S
Site 118
T1232
E
A
L
L
E
A
K
T
N
E
L
I
N
I
S
Site 119
S1248
S
K
T
N
A
I
L
S
R
I
S
H
C
Q
H
Site 120
T1258
S
H
C
Q
H
R
T
T
K
V
K
E
A
L
L
Site 121
T1282
E
A
Q
L
R
Q
L
T
E
E
Q
N
T
L
N
Site 122
T1287
Q
L
T
E
E
Q
N
T
L
N
I
S
F
Q
Q
Site 123
S1291
E
Q
N
T
L
N
I
S
F
Q
Q
A
T
H
Q
Site 124
S1308
E
K
E
N
Q
I
K
S
M
K
A
D
I
E
S
Site 125
T1318
A
D
I
E
S
L
V
T
E
K
E
A
L
Q
K
Site 126
S1339
Q
A
A
S
E
K
E
S
C
I
T
Q
L
K
K
Site 127
T1342
S
E
K
E
S
C
I
T
Q
L
K
K
E
L
S
Site 128
S1349
T
Q
L
K
K
E
L
S
E
N
I
N
A
V
T
Site 129
S1371
E
K
K
V
E
I
S
S
L
S
K
Q
L
T
D
Site 130
S1373
K
V
E
I
S
S
L
S
K
Q
L
T
D
L
N
Site 131
T1377
S
S
L
S
K
Q
L
T
D
L
N
V
Q
L
Q
Site 132
S1386
L
N
V
Q
L
Q
N
S
I
S
L
S
E
K
E
Site 133
S1388
V
Q
L
Q
N
S
I
S
L
S
E
K
E
A
A
Site 134
S1390
L
Q
N
S
I
S
L
S
E
K
E
A
A
I
S
Site 135
S1397
S
E
K
E
A
A
I
S
S
L
R
K
Q
Y
D
Site 136
S1398
E
K
E
A
A
I
S
S
L
R
K
Q
Y
D
E
Site 137
Y1403
I
S
S
L
R
K
Q
Y
D
E
E
K
C
E
L
Site 138
S1418
L
D
Q
V
Q
D
L
S
F
K
V
D
T
L
S
Site 139
T1423
D
L
S
F
K
V
D
T
L
S
K
E
K
I
S
Site 140
S1425
S
F
K
V
D
T
L
S
K
E
K
I
S
A
L
Site 141
T1454
K
K
A
Q
S
R
F
T
Q
H
Q
N
T
V
K
Site 142
T1459
R
F
T
Q
H
Q
N
T
V
K
E
L
Q
I
Q
Site 143
S1471
Q
I
Q
L
E
L
K
S
K
E
A
Y
E
K
D
Site 144
Y1475
E
L
K
S
K
E
A
Y
E
K
D
E
Q
I
N
Site 145
S1507
G
E
M
E
D
D
K
S
K
M
E
K
K
E
S
Site 146
S1514
S
K
M
E
K
K
E
S
N
L
E
T
E
L
K
Site 147
T1518
K
K
E
S
N
L
E
T
E
L
K
S
Q
T
A
Site 148
S1522
N
L
E
T
E
L
K
S
Q
T
A
R
I
M
E
Site 149
T1535
M
E
L
E
D
H
I
T
Q
K
T
I
E
I
E
Site 150
T1538
E
D
H
I
T
Q
K
T
I
E
I
E
S
L
N
Site 151
S1543
Q
K
T
I
E
I
E
S
L
N
E
V
L
K
N
Site 152
Y1551
L
N
E
V
L
K
N
Y
N
Q
Q
K
D
I
E
Site 153
T1589
E
A
E
E
K
I
L
T
L
E
N
Q
V
Y
S
Site 154
Y1595
L
T
L
E
N
Q
V
Y
S
M
K
A
E
L
E
Site 155
S1614
E
L
E
H
V
N
L
S
V
K
S
K
E
E
E
Site 156
S1617
H
V
N
L
S
V
K
S
K
E
E
E
L
K
A
Site 157
S1631
A
L
E
D
R
L
E
S
E
S
A
A
K
L
A
Site 158
S1633
E
D
R
L
E
S
E
S
A
A
K
L
A
E
L
Site 159
S1657
A
I
K
K
Q
L
L
S
Q
M
E
E
K
E
E
Site 160
S1672
Q
Y
K
K
G
T
E
S
H
L
S
E
L
N
T
Site 161
S1675
K
G
T
E
S
H
L
S
E
L
N
T
K
L
Q
Site 162
T1679
S
H
L
S
E
L
N
T
K
L
Q
E
R
E
R
Site 163
S1697
I
L
E
E
K
L
K
S
V
E
S
S
Q
S
E
Site 164
S1700
E
K
L
K
S
V
E
S
S
Q
S
E
T
L
I
Site 165
S1701
K
L
K
S
V
E
S
S
Q
S
E
T
L
I
V
Site 166
S1703
K
S
V
E
S
S
Q
S
E
T
L
I
V
P
R
Site 167
T1705
V
E
S
S
Q
S
E
T
L
I
V
P
R
S
A
Site 168
S1726
T
E
Q
E
E
A
D
S
Q
G
C
V
Q
K
T
Site 169
T1733
S
Q
G
C
V
Q
K
T
Y
E
E
K
I
S
V
Site 170
Y1734
Q
G
C
V
Q
K
T
Y
E
E
K
I
S
V
L
Site 171
S1739
K
T
Y
E
E
K
I
S
V
L
Q
R
N
L
T
Site 172
T1746
S
V
L
Q
R
N
L
T
E
K
E
K
L
L
Q
Site 173
T1762
V
G
Q
E
K
E
E
T
V
S
S
H
F
E
M
Site 174
S1764
Q
E
K
E
E
T
V
S
S
H
F
E
M
R
C
Site 175
S1765
E
K
E
E
T
V
S
S
H
F
E
M
R
C
Q
Site 176
Y1773
H
F
E
M
R
C
Q
Y
Q
E
R
L
I
K
L
Site 177
S1792
A
K
Q
H
E
D
Q
S
M
I
G
H
L
Q
E
Site 178
S1809
E
E
K
N
K
K
Y
S
L
I
V
A
Q
H
V
Site 179
T1841
V
F
D
D
V
Q
K
T
L
Q
E
K
E
L
T
Site 180
T1848
T
L
Q
E
K
E
L
T
C
Q
I
L
E
Q
K
Site 181
S1861
Q
K
I
K
E
L
D
S
C
L
V
R
Q
K
E
Site 182
S1879
V
E
M
E
E
L
T
S
K
Y
E
K
L
Q
A
Site 183
T1904
T
E
L
L
E
E
N
T
E
E
K
S
K
S
H
Site 184
S1910
N
T
E
E
K
S
K
S
H
L
V
Q
P
K
L
Site 185
S1919
L
V
Q
P
K
L
L
S
N
M
E
A
Q
H
N
Site 186
T2015
K
E
Q
E
L
E
M
T
I
K
E
T
I
N
K
Site 187
S2033
V
E
A
E
L
L
E
S
H
Q
E
E
T
N
Q
Site 188
Y2059
L
K
R
T
A
K
R
Y
E
E
I
L
D
A
R
Site 189
T2071
D
A
R
E
E
E
M
T
A
K
V
R
D
L
Q
Site 190
T2079
A
K
V
R
D
L
Q
T
Q
L
E
E
L
Q
K
Site 191
T2110
V
T
I
M
E
L
Q
T
Q
L
A
Q
K
T
T
Site 192
T2116
Q
T
Q
L
A
Q
K
T
T
L
I
S
D
S
K
Site 193
T2117
T
Q
L
A
Q
K
T
T
L
I
S
D
S
K
L
Site 194
S2120
A
Q
K
T
T
L
I
S
D
S
K
L
K
E
Q
Site 195
Y2143
L
E
D
R
L
K
K
Y
E
K
N
V
Y
A
T
Site 196
Y2148
K
K
Y
E
K
N
V
Y
A
T
T
V
G
T
P
Site 197
T2150
Y
E
K
N
V
Y
A
T
T
V
G
T
P
Y
K
Site 198
T2154
V
Y
A
T
T
V
G
T
P
Y
K
G
G
N
L
Site 199
Y2156
A
T
T
V
G
T
P
Y
K
G
G
N
L
Y
H
Site 200
Y2162
P
Y
K
G
G
N
L
Y
H
T
D
V
S
L
F
Site 201
T2164
K
G
G
N
L
Y
H
T
D
V
S
L
F
G
E
Site 202
S2167
N
L
Y
H
T
D
V
S
L
F
G
E
P
T
E
Site 203
T2173
V
S
L
F
G
E
P
T
E
F
E
Y
L
R
K
Site 204
Y2177
G
E
P
T
E
F
E
Y
L
R
K
V
L
F
E
Site 205
Y2185
L
R
K
V
L
F
E
Y
M
M
G
R
E
T
K
Site 206
T2191
E
Y
M
M
G
R
E
T
K
T
M
A
K
V
I
Site 207
T2209
L
K
F
P
D
D
Q
T
Q
K
I
L
E
R
E
Site 208
T2223
E
D
A
R
L
M
F
T
S
P
R
S
G
I
F
Site 209
S2224
D
A
R
L
M
F
T
S
P
R
S
G
I
F
_
Legend
Confirmed in mammals
Confirmed in related proteins or other species
Predicted by Kinexus P-Site Prediction algorithm
No data/link available
Link available
Products available
2019 Kinexus Bioinformatics Corporation